Four-time consecutive U.S. Junior tournament champ Ryoma Echizen comes to Seishun Academy to further his reign as The Prince of Tennis. His skill is matched only by his attitude--irking some but impressing all as he leads his team to the Nationals and beyond!

As the road to the Nationals narrows, Seishun Academy squares off with St. Rudolph. Through the maneuverings of Hajime, Saint Rudolph's cunning manager, Ryoma gets matched up with Yuta-known to many as Lefty Killer. With an unorthodox game laced with reverse spins and angles, Yuta's game ignites against the seemingly inexperienced-against-lefties Ryoma. Meanwhile, the Seishun doubles tandem of Eiji and Shuichiro assume the Australian formation to break a tie. Is this a desperate attempt to win, or a well-calculated strategy?